Brownheaded leafrollers - Mortality
No comprehensive analysis has been carried out of the importance and role of mortality factors affecting brownheaded leafroller populations in apple orchards in New Zealand. The following Table summarises the limited results of several studies on apples.
Mortality factor Mortality Causes
Unfulfilled fecundity ?Unknown Mainly weather, host plant, larval density
Infertility <1%  
Inviability 2.5% Weather
Egg predation/parasitism 3-12% Predatory bugs, Anystis baccarum, Trichogramma spp.
Predation and losses of young larvae (between hatching and larval establishment on apple trees) 88-94% Spiders, earwigs, predatory bugs, Anystis baccarum, emigration, weather, birds
Parasitism of larvae, all species ?Unknown Complex of parasitoids
Larval/pupal parasitism ?Unknown Pales spp. and complex of introduced parasitoids.
Emigration from orchards - probably minor
Immigration to orchards - probably major
  High populations on nearby host plants causing immigration into low resident density in orchards